I agree with OWD that this is not a Jones Underlever. The operating lever is like that of the Jones but this appears to be a single bite locking lug while the Jones patent screw grip was for double opposing bite locking lugs.
You can see the two bites in this photo of my Lang:

Your gun should be fall between 1870 and 1875 - my Lang s/n 32XX letters to March 6, 1866. Don Masters book "Atkin, Grant & Lang" lists serial number groups as follows:
1865 - #2882
1870 - #3851
1875 - #5125
Don Masters lists 1875 as the year when the name was changed from Joseph Lang to Joseph Lang & Sons.
I don't know the current cost but you can get a letter from AG&L by writing or emailing Francis Lovel the current owner.
Atkin, Grant & Lang
Windmill Road, Markyate, St Albans,
Hertfordshire AL3 8BP, England
francis@atkingrantandlang.com
www.atkingrantandlang.comTel +44 (0)1993 864 949
Fax +44 (0)7812 375 781